01. ∀x[Qx↔[∃y[[Rxy]∧¬[x=y]]]] premise
02. ∀xQx premise
03. ∃x∃y∀z[[z=x]∨[z=y]] premise
04. ∃y∀z[[z=a]∨[z=y]] assumption
05. ∀z[[z=a]∨[z=b]] assumption
06. Rcd assumption
07. ¬[c=d] assumption
08. ∀x[Qx↔[∃y[[Rxy]∧¬[x=y]]]] restate 01
09. Qd↔[∃y[[Rdy]∧¬[d=y]]] ∀elim 08
10. ∀xQx restate 02
11. Qd ∀elim 10
12. ∃y[[Rdy]∧¬[d=y]] ↔elim 09 11
13. [Rde]∧¬[d=e] assumption
14. e=c assumption
15. [Rde]∧¬[d=e] restate 13
16. [Rdc]∧¬[d=c] para 15 14
17. Rdc ∧elim 16
18. [e=c]→[Rdc] →intro 19 17
19. ¬[e=c] assumption
20. ∀z[[z=a]∨[z=b]] restate 05
21. [c=a]∨[c=b] ∀elim 20
22. [d=a]∨[d=b] ∀elim 20
23. [e=a]∨[e=b] ∀elim 20
24. [Rde]∧¬[d=e] restate 13
25. ¬[d=e] ∧elim 24
26. e=a assumption
27. ¬[d=e] restate 25
28. ¬[d=a] para 27 26
29. [d=a]∨[d=b] restate 22
30. d=b ∨elim 29 28
31. ¬[e=c] restate 19
32. ¬[a=c] para 31 26
33. ¬[c=a] flip 32
34. [c=a]∨[c=b] restate 21
35. c=b ∨elim 34 33
36. ¬[c=d] restate 07
37. c=d para 35 30
38. ⊥ 36 37
39. ¬[e=a] ¬intro 26 38
40. e=b ∨elim 23 39
41. ¬[d=e] restate 25
42. ¬[d=b] para 41 40
43. d=a ∨elim 22 42
44. ¬[c=d] restate 07
45. ¬[c=a] para 44 43
46. c=b ∨elim 21 45
47. e=c para 40 46
48. ⊥ 47 19
49. ¬¬[e=c] ¬intro 19 48
50. e=c ¬¬elim 49
51. Rdc MP 18 50
52. Rdc ∃ 12 13-51
53. ¬[c=d]→[Rdc] →intro 07 52
54. c=d assumption
55. Rcd restate 06
56. Rcc para 55 54
57. Rdc para 56 54
58. [c=d]→[Rdc] →intro 54 57
59. [c=d]∨¬[c=d] LEM
60. Rdc ∨elim 58 53 59
61. [Rcd]→[Rdc] →intro 06 60
62. ∀y[[Rcy]→[Ryc]] ∀intro 61
63. ∀x∀y[[Rxy]→[Ryx]] ∀intro 62
64. ∀x∀y[[Rxy]→[Ryx]] ∃elim 04 05-63
65. ∀x∀y[[Rxy]→[Ryx]] ∃elim 03 04-64
66. ∀y[[Ray]→[Rya]] ∀elim 65
67. [Rab]→[Rba] ∀elim 66
68. ∀y[[Rby]→[Ryb]] ∀elim 65
69. [Rba]→[Rab] ∀elim 68
70. [Rab]↔[Rba] ↔intro 67 69
71. ∀y[[Ray]↔[Rya]] ∀intro 70
72. ∀x∀y[[Rxy]↔[Ryx]] ∀intro 71
Synopsis of proof:
We let $Rcd$. Then, using the premise $\forall x \exists y [Rxy \land \neg x = y]$, we conclude $Rde$. Then, we argue that $e$ must be equal to $c$, using $\forall z[z = a \lor z = b]$. Then, we conclude that $Rcd \implies Rdc$. Using symmetry, we conclude that $Rdc \implies Rcd$, hence $Rcd \iff Rdc$. Then, we generalize to $\forall x \forall y[Rxy \iff Ryx]$.