The meaning of the LHS is as follows: we first define $f: \Bbb{R}^2 \to \Bbb{R}$ to be the function $f:= \dfrac{\partial T}{\partial x}$. In other words, for every $(\alpha, \beta) \in \Bbb{R}^2$,
\begin{align}
f(\alpha, \beta) := \dfrac{\partial T}{\partial x}(\alpha, \beta)
\end{align}
So, we now want to express $\dfrac{\partial f}{\partial y}(0,y_0)$ in a different way. Well, by definition,
\begin{align}
\dfrac{\partial f}{\partial y}(0,y_0) &:= \dfrac{d}{dy} \bigg|_{y=y_0}\bigg( y \mapsto f(0,y)\bigg) \tag{$*$}\\
&:= \dfrac{d}{dy} \bigg|_{y=y_0}\bigg(y \mapsto \dfrac{\partial T}{\partial x}(0, y) \bigg)
\end{align}
So, really, this equality is true by definition of a partial derivative. Now, at this point, you might find my first equality a little suspicious and wonder if it is really true by definition. Well, ok depending on how you look at it, it's either a definition or a simple theorem with a one-line-proof (so really, I just consider it a definition).
You're probably more familiar with this definition:
Let $f: \Bbb{R}^2 \to \Bbb{R}$ be a function, and $(\alpha, \beta) \in \Bbb{R}^2$ be a point. If the limit
\begin{align}
\lim_{h \to 0} \dfrac{f(\alpha, \beta + h) - f(\alpha, \beta)}{h}
\end{align}
exists, we denote it as $\dfrac{\partial f}{\partial y}(\alpha, \beta)$, or $(\partial_2f)(\alpha, \beta)$, and call it the partial derivative of $f$ with respect to the second variable, evaluated at the point $(\alpha, \beta)$.
But if you look carefully, what is this definition saying? Well, the existence of that limit is precisely equivalent to the differentiability of the function $y \mapsto f(\alpha, y)$ at the point $\beta$. In other words, the partial derivative (if it exists) is exactly equal to any of the following:
\begin{align}
\dfrac{\partial f}{\partial y}(\alpha, \beta) &:= \bigg( y \mapsto f(\alpha, y)\bigg)'(\beta) \\
&\equiv \dfrac{d}{dy} \bigg|_{y = \beta} \bigg( y \mapsto f(\alpha, y)\bigg)\\
&= \dfrac{d}{ds} \bigg|_{s = 0} \bigg( s \mapsto f(\alpha, \beta + s)\bigg)\\
&= \dfrac{d}{ds} \bigg|_{s = 0} \bigg( s \mapsto f\left((\alpha, \beta) + s e_2 \right)\bigg)
\end{align}
where the $\equiv$ means "same thing expressed in different notation", and the third line is true by a simple application of the single variable chain rule (or just look at the limit definition, it's trivially true), and $e_2 = (0,1)$.
BTW, you're perhaps right to introduce a new function $\mathcal{T}$ to avoid any abuse of notation, but I didn't feel like coming up with new names for each new function I defined along the way, which is why I used the stopped arrow $\mapsto$ notation to indicate precisely what the function is, and I was also careful to distinguish between the bound and free variables, and a function vs the function's values at a point so hopefully it's clear exactly how to read the above notation.